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Chrysellampus Semenov-Tian-Shanskij, 1932: 5. Type species: Ellampus heros Semenov, 1892, by original designation.
Parellampus Semenov-Tian-Shanskij, 1932: 7. Type species: Parellampus praeteritorum Semenov, 1932, by original designation. Syn. nov.
Chrysellampus Semenov: Semenov-Tian-Shanskij 1967: 119; Nikol'skaya 1978: 63 (part.).
Omalus (Chrysellampus) Panzer, 1801: Linsenmaier 1959: 22, 1997: 249 (part.), 1999: 22, 23 (part.); Martynova & Fateryga 2014: 14.
Omalus (Philoctetes) Panzer: Arens 2014: 570 (part.).

Systematics. The genus Chrysellampus Semenov, 1932 was traditionally considered as a valid genus (Tsuneki 1948, 1950, 1953a, 1953b; Semenov 1967; Nikol'skaya 1978) or valid subgenus (Linsenmaier 1959, 1997; Martynova & Fateryga 2014). Kimsey & Bohart (1991) proposed a new systematic classification of genera closely related to Omalus Panzer, 1801 and (1991: 251) synonymised Chrysellampus Semenov, 1932 with Philoctetes Abeille de Perrin, 1879. Later Linsenmaier (1997) revalidated Chrysellampus Semenov, 1932 as subgenus.

Description. Chrysellampus Semenov, 1932 was synonymised with Philoctetes Abeille de Perrin, 1879 by Kimsey & Bohart (1991), but it can be easily separated by the following characteristics: habitus subcylindrical and elongate (stocky and roundish in Philoctetes) (Martynova & Fateryga 2014: Fig. 14); F2–F11 flattened and dilated (Fig. 3) (cylindrical and not dilated in Philoctetes); MS bisected or nearly so by curved genal carina (Figs 2A, 3, 5, 8) (MS not bisected and genal carina not curved in Philoctetes); genal carina well developed and sharp from lower margin of temple to MS (Figs 3, 5, 8) (faint or not well developed in Philoctetes); pronotum with large and dense punctures (Figs 4A–4F, 6A–6F, 8) (smooth to sparsely punctate in Philoctetes); mesoscutum with large punctures clumped along notauli (C. sculpticollis and C. medanae) or covering entire surface (punctures only clumped along notauli or more evenly distributed, but anyway gathered together toward notauli in Philoctetes); if punctures on mesoscutum are clumped along notauli, then punctuation on pronotum is dense with large punctures (smooth to sparsely punctate in Philoctetes); metanotum round, with exception of C. praeteritorum (Semenov, 1932) (usually conical to mucronate in Philoctetes); medial vein weakly curved (strongly arched in Philoctetes); the distance between posterior margin of anterior declivity of T1 and posterior margin of T1 as long as or longer than mesoscutellum (Fig. 2C) (very short, 0.2–0.5 × as long as in Philoctetes (Fig. 2B)); apex of T3 with median notch, with tooth at each side (with or without median notch in Philoctetes); head and mesosoma characterized by colliculate sculpture (Harris 1979), a reticulate microsculpture continuously set with granulations on the intervals among punctures (Figs 1A, 1B) (polished in Philoctetes); tarsal claw with five teeth (two to four in Philoctetes).

FIGURE 1. Chrysellampus sculpticollis (Abeille de Perrin), ♀. A. Punctuation and colliculate sculpture of mesoscutum over tegula margin, 150x; B. Punctuation and colliculate sculpture of mesoscutum over tegula margin, close up, 300x; C. Left apical tooth with rim, 500x. (Agnoli & Rosa 2011).

FIGURE 2. A. Chrysellampus sculpticollis (Abeille de Perrin), ♂. Head, lateral view. Arrow pointing curved genal carina bisecting MS.

FIGURE 2. B. Philoctetes sp., metasoma, dorsal view. Arrow delimiting the distance between posterior margin of anterior declivity and posterior margin of T1; C. Chrysellampus sp., metasoma, dorsal view. Arrow delimiting the distance between posterior margin of anterior declivity and posterior margin of T1.

Species included. At present, the genus Chrysellampus Semenov includes ten rare and homogeneous species: C. sculpticollis (Abeille de Perrin, 1878), C. medanae (du Buysson in Magretti, 1890), C. heros (Semenov, 1892), C. pici (du Buysson, 1900) (=C. nigromaculatus Linsenmaier, 1997), C. harmandi (Buysson, 1903), C. praeteritorum (Semenov, 1932), comb. nov., C. duplipunctatus Tsuneki, 1948, C. tatianae Semenov, 1967, C. obtusidentibus Rosa, Wei & Xu, sp. nov., and C. proximocellis Rosa, Wei & Xu, sp. nov.

Biology. Chrysellampus are known as nest parasite of Crabronidae (Martynova & Fateryga 2014). Martynova & Fateryga (2014) gave description of larva and cocoon of C. sculpticollis, and its feeding habits and interactions with host, Psenulus fuscipennis (Dahlbom).

Distribution. Chrysellampus Semenov is known from Palaearctic and Oriental Regions (China: Yunnan).

From: Rosa P., Wei N.-S. & Xu Z.-F., 2015 - Revalidation of genus Chrysellampus Semenov, 1932, with description of two new species from China (Hymenoptera, Chrysididae). Zootaxa 4034 (1): 148–160.

European Species

Pentachrysis Lichtenstein, 1876: 227.
Type species: Chrysis amoena Eversmann 1858 [= Pentachrysis amoena (Eversmann, 1858)], by subsequent designation of Ashmead 1902: 226.

Distribution: Ethiopian, Oriental and Palaearctic Region.

Gonodontochrysis Semenov-Tian-Shanskij, 1954a (as subgenus of Chrysis Linnaeus, 1761): 120. Unavailable name.
Chrysis (Chrysis) pulchella group: Linsenmaier 1959a: 93 (key), 103 (diagnosis).
Chrysis pulchella group: Kimsey & Bohart 1991: 322 (key), 331 (Fig. 107o), 336 (Fig. 110h), 358 (diagnosis, discussion). Farhad et al. 2019: 1006 (diagnosis).
Chrysis zaravshanica group: Tarbinsky 2002: 23 (description).

Some of the species included in this genus were previously included in Gonodontochrysis Semenov-TianShanskij, 1954. Linsenmaier (1959) and Kimsey & Bohart (1991) included all members of Morphochrysis in the Chrysis pulchella species-group (Fig. 1). Tarbinsky (2002) described the Chrysis zaravshanica species-group based on a member of this genus (Chrysis personata Semenov, 1967 = Chrysis zaravshanica Tarbinsky, 2002 syn. nov.).

Description. Species of medium to large size, with head distinctly larger than pronotum; first flagellomere elongate in both sexes (length/width ratio 2.5–3.2); scapal basin medially polished or slightly corrugated in the upper part, occasionally finely punctate in females; face usually finely punctate and covered by dense, adpressed whitish pubescence in males; transverse frontal carina strongly developed, broadly M-like, sometimes with distinct rami almost encircling anterior ocellus or delimiting anterior ocellar area; malar space as long as 1.0–1.5 × anterior ocellus diameter; radial cell open, with fore-wing radial sector short, 0.5–2.5 × anterior ocellus diameter away from wing margin; second metasomal tergum with weak to moderate longitudinal medial carina; third metasomal tergum with distinct pit row; lateral edges of third tergum with a small tooth, or angle, at or beyond the middle, followed by a slight concavity, more or less marked depending on species; third tergum apicomedially usually biconvex, rarely convex (Morphochrysis diadema) to nearly straight (Morphochrysis atechka and Morphochrysis intercurra) (Fig. 2); black spots on second sternum oval or rectangular, in some species close to each other or completely fused medially and covering large part of the segment (e.g. Morphochrysis pulchella and Morphochrysis calimorpha); male eighth sternite quadrangular, apically broad; male genital capsule with slender and elongate gonocoxa, apically curved; male and female internal segments unusually round shaped (Fig. 3). Finally, this genus is supported by molecular phylogenetic analyses as a distinct clade (Pauli et al. 2019).

Figure 1. Morphochrysis gen. nov., habitus, dorsal view. A. M. asahinai, ♀; B. M. andradei, ♀; C. M. atechka, ♀; D. M. atechka, ♂; E. M. pulchella, ♂; F. M. pulchella, ♀; G. M. dives, ♂; H. M. dives, ♀; I. M. personata, ♂; J. M. rubicunda, ♀; K. M. trisinuata, ♀; L. M. turceyana, ♀.

Figure 2. Morphochrysis gen. nov., third metasomal tergum, postero-lateral view. A. M. personata, ♂; B. M. pulchella, ♀; C. M. diadema, ♂; D. M. atechka, ♀; E. M. asahinai, ♀; F. M. urakensis, ♂; G. M. mosulensis, ♀; H. M. tedshensis, ♀.

Figure 3. Morphochrysis gen. nov., fifth and sixth female metasomal terga. A–B. M. pulchella; C–D. M. calimorpha; E–F. M. goetheana.

Colpopyga Semenov-Tian-Shanskij, 1954: 137.
Colopyga: Kimsey & Bohart 1991: 181, incorrect subsequent spelling.
Type species: Hedychrum flavipes Eversmann, 1858, by original designation.

Systematics. Colpopyga Semenov-Tian-Shanskij, 1954, is a small genus of cuckoo wasps and was synonymised with Hedychridium Abeille de Perrin, 1878 by Linsenmaier (1959) and its species were included in the H. flavipes species group (Linsenmaier 1968). Noskiewicz & Lorencowa (1963) and other authors considered Colpopyga a valid genus based on the modified structure of the female metasoma, in particular of the cone-shaped T3 and the modified structure of the internal sterna and terga. Finally, molecular data published by Pauli et al. (2019) placed Colpopyga on a monophyletic clade with a clear affinity with the genus Holopyga Dahlbom, 1854 rather than Hedychridium.

Diagnosis. The genus Colpopyga is characterised by the female metasoma highly modified and flattened in lateral view; the T3 is elongate, cone-shaped; the structure of internal terga and sterna is modified, highly elongated; the male T3 is dimorphic, evenly rounded; the male genitalia are weakly pigmented; in both sexes, the flagellomeres are elongate, cylindrical, with F1 at least twice as long as broad; the propodeal tooth (= posterior propodeal projection) is spiniform; the legs are largely or entirely non-metallic, yellow; the tegulae are non-metallic brown; the apical margin of T3 has a narrow hyaline-brownish rim, usually medially emarginate; the transverse frontal carina is faint; the face has sparse erect setae; the malar spaces are less than 1 MOD; the scapal basin is shallow, medially cross-ridged; the mandible is tridentate; the inner surface of the mid and hind tibia is without pits; the medial vein of forewing is arched; the tarsal claw bears a single submedian, perpendicular tooth.

Fig. 1. Habitus, females, dorsal view. A, Colpopyga flavipes flavipes (Eversmann), Italy; B, C. flavipes rugulosa (Linsenmaier), Cyprus; C, C. temperata (Linsenmaier), holotype, Morocco; D, C. auriventris (Mercet), Spain; E, C. nesterovi sp. nov., paratype, Kazakhstan. Scale bar: 1.0 mm.

Fig. 2. Habitus, males, dorsal view. A, Colpopyga flavipes flavipes (Eversmann), Italy; B, C. flavipes rugulosa (Linsenmaier), paratype, Iran; C, C. temperata (Linsenmaier), Morocco; D, C. auriventris (Mercet), Spain; E, C. nesterovi sp. nov., paratype, Kazakhstan. Scale bar: 1.0 mm.

Fig. 3. Habitus, females, lateral view. A, Colpopyga flavipes flavipes (Eversmann), Italy; B, C. flavipes rugulosa (Linsenmaier), Cyprus; C, C. temperata (Linsenmaier), paratype, Tunisia; D, C. auriventris (Mercet), Spain; E, C. nesterovi sp. nov., paratype, Kazakhstan. Scale bar: 1.0 mm.

Fig. 4. T3, dorso-lateral view. A, Colpopyga auriventris (Mercet), female, Spain; B, C. nesterovi sp. nov., female, holotype, Kazakhstan; C, C. nesterovi sp. nov., male, paratype, Kazakhstan; D, C. flavipes flavipes (Eversmann), female, Italy; E, C. flavipes rugulosa (Linsenmaier), female, Cyprus; F, C. temperata (Linsenmaier), female, holotype, Morocco; G, C. temperata (Linsenmaier), male, Tunisia; H, C. flavipes cyanomaculata (Trautmann), male, lectotype, Tunisia; I, Hedychridium elongatum Linsenmaier, holotype, Morocco.